Mine's far enough back I don't rightly remember. I THINK it was something a friend came up with for something or other, and I took it and used it for a handle. Since that time in 3rd grade or so, it's become my online handle for EVERYTHING online (seriously, google it, and there will be about an entire page of accounts made by me there), as well as my XBL handle and my first original D&D character. (In progress. The origin story was pretty awesome, if cliche ridden)
Everyone I know online knows me by this name (I know at least one (annoying) person who calls me that irl, since we know others with my actual name), and I've got a blog, a Youtube channel, and a twitter, all with this name, so hopefully one of these days it'll be marginally famous!
Mine is based off a long story I will not go into detail for, but Luna is Moon en espanol, and nite is, well, night in short. Simply, but the story of the night theme to my name goes on to a list I hate explaining, so I'll leave it there.
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A variation on the spelling of Ikarus that I've used forever. It's just one of 3 of my most common usernames.
Edit: The other two being Delpheus and Suicidal Ferret. Delpheus is a character from Fire Warrior; a character with the ability to see the future. Using this ability, he told his squadmates not to fight, that the greater threat was to be seen. They didn't believe him until he died. It's the same with my friends; they never listen until it's too late.
Suicidal Ferret is based on a variety of things. I love ferrets, my gaming style can be decribed as suicidal (maybe fanatical is a better word in hind sight), and lastly, I've had issues with the will to live.
My friends and I all love Red Vs Blue and we all kind of match one of the characters from the series in personalities. My best friend acts just like Caboose, my little brother is Grif and I was christened as Tex. The Aussie part was just added to the front because Tex was taken and I am Australian.
My name's Orin, and there aren't too many characters who share my name. I used to play the Mage Knight collectible miniatures game, and they came out with a hero called Lord Oren. I was pretty happy... except my friend wouldn't frickin' trade hers to me XD