i have no problem personally with a handshake, cant imagine anyone would. a hug is completely normal, so long as it dosent turn into dry humping but that goes for everyone, not just hetero guys (unless its porn. then anything goes).
a kiss on the cheek/forehead is also completely normal. i mean, i wouldnt do it to my guy friends, but thats because im a hugger, and i think i would rather not be labeled gay when im only a freshman (because SOMEONE is going to think that, and say it very loudly. fuck teenagers. seriously. soo many of them suck)
hand holding... is suggestive. i can see little kids doing it, or really good female friends of any age (have actually seen it), but when i picture two grown guys holding hands, just seems too suggestive since all the men i know are not really prone to prolonged contact. keep it to a handshake or a hug they do. so i would draw the line at hand holding, for it seems out of place to me.
but hell, if you got a good guy friend, and he is hurting, cuddling is even okay in some situations like that. so yeah, my lines are a bit different than the poll's.